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Yamaha's flagship 450cc four-stroke motocross racer. Built for 450-class racers and experienced adult riders.
| 1999 | 113.4 kg |
| 2000 | 998 mm |
| 2001 | 998 mm | US$5,999 |
| 2002 | 998 mm | US$5,999 |
| 2003 | 998 mm | US$6,299 |
| 2004 | 100.7 kg | 996 mm | US$6,299 |
| 2005 | 99.8 kg | 996 mm | US$6,399 |
| 2006 | 99.8 kg | 986 mm | US$6,899 |
| 2007 | 99.8 kg | 988 mm | US$6,999 |
| 2008 | 99.3 kg | 988 mm | US$7,099 |
| 2009 | 988 mm | US$7,399 |
| 2010 | 998 mm | US$7,990 |
| 2011 | 998 mm | US$8,150 |
| 2012 | 1,001 mm | US$8,350 |
| 2013 | 1,001 mm | US$8,490 |
| 2014 | 975 mm | US$8,490 |
| 2015 | 975 mm | US$8,590 |
| 2016 | 965 mm | US$8,590 |
| 2017 | 965 mm | US$8,699 |
| 2018 | 980 mm | US$9,199 |
| 2019 | 980 mm | US$9,299 |
| 2020 | 965 mm | US$9,399 |
| 2021 | 965 mm | US$9,399 |
| 2022 | 965 mm | US$9,599 |
| 2023 | 965 mm | US$9,899 |
| 2024 | 965 mm | US$9,999 |
| 2025 | 965 mm | US$9,999 |
| 2026 | 965 mm | US$10,299 |
| 2027 | 965 mm | US$10,299 |